Using the molecular clock method, it was possible to determine the age of the major branches of the orchid family. This also confirmed that the subfamily Vanilloideae is a branch at the basal dichotomy of the monandrous orchids, and must have evolved very early in the evolution of the family. Since this subfamily occurs worldwide in tropical and subtropical regions, from tropical America to tropical Asia, New Guinea and West Africa, and the continents began to split about 100 million years ago, significant biotic exchange must have occurred after this split (since the age of Vanilla is estimated at 60 to 70 million years).
Special thanks to David Banks of Australia, Christian Kirsch; Dr. Rudolph Jenny, Peter O'Byrne***, Eric Hagsater***, Lisa Thoerle***, Mario Blanco***, Mark Whitten***, Adam Karremans, Hakan Halander****; Daniel Jimenez***, André Schuiteman***, Dr. E.F. de Vogel, Jaap Vermeulen, Rick Barry, Dr. Guido Braeme, Rogier Van Vugt; Rene Dishong, Tom Ballinger, Mac & Helen Rivenback; Dr Rubin Salueda; Rafael Govaerts***, Padre Pedro Ortiz, Dr Leslie Garay, Vena Read, Charles Lamb, Donald Tan of Malaysia, Joseph Dougherty, Dalton Holland Baptista, Rick Cirino; Prem Subrahmanyam, ROMAN MARUSKA Czech Republic, Michael Coronado, Alejandro Taborda, Greg Butler and Oak Hill Gardens, Bill Bergstrom and Bergstrom Orchids, Donna Nash, Oliver Lenhard, D. E Vermeullen, Americo Docha Neto, Richard Korber, Carl Withner, Mark Nir, Ekkehard Schwadtke; Dr Eric Christenson, Dave Alford, Igor Zhuravlev, Tony Watkinson, David Hunt, Jean Marie Vanderwinden, Glen Ladnier, Jeffrey W. Tucker, Jorge Alejandro Paulete Scaglia, Scott McGregor, Wilella Stimmell, Eric Thiessen, Geert Volckaert, George and Kathy Norris***, David Alford, Eric Hunt, Peter O'Byrne, Allen Black, Helen Milner, Al Pickrel, Greg Riemer, Marcia Whitmore, Phillipe Musschoot, Wilford Neptune, Jean-Claude George, Tony Walsh, Greg and Kerri Steenbeeke and their Orkology Kreations Website, Jim Hamilton, Robert Fuchs and RF Orchids, Gene Monier and JEM Orchids, Jerry Bolce, Andy's Orchids, Stephen Jones, First Ray Orchids, Robert Weyman Bussey, JEM Orchids, Carlos Hajek and his Peruvian Orchid Page, Bill Morden, David Morris, Noble Bashor, Patricia Harding, Nina Rach, David Haelterman; Craig Cooper, Rocky Giovinazzo, David Friedman, Bill Pinnix, Troy C. Meyers, Linda's Orchid Page, Pierre-Alain Darlet*, Guy Cantor, Tropical Orchid Farm in Maui, H&R Nursuries, Lois Greer, Ichiro "Haru" Ohsaka and his Bulbophyllum Page, Mauro Rosim from Brazil, A World Of Orchids, Bruce Norris of Canada, Nick Doe, Neville De La Rue, Nelson Barbosa Machado Neto, Jeff Aguillon, Dale and Deni Borders, Eka Mulja Tjipta, Rick Barry, Manfred Schmucker, Uri Baruk, Mike and Candy Joehrendt, Dan and Marla Nikirk, Walter Orchard, Christer Carlson; Guillerrmo Angulo, M. Max, Paula Vagner, Tennis Maynard, Greg Allikas, Malcolm Thomas, David Morris, Jacques Deschenes, Lourens Grobler; Gary Yong Gee; José Castaño Hernández; Judd and David Janvrin* for their photo and text contributions.
The seeds are generally almost microscopic and very numerous, in some species over a million per capsule. After ripening, they blow off like dust particles or spores. They lack endosperm and must enter symbiotic relationships with various mycorrhizal basidiomyceteous fungi that provide them the necessary nutrients to germinate, so all orchid species are mycoheterotrophic during germination and reliant upon fungi to complete their lifecycles.